Ian Gilby
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Phone: 480-965-3807
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ISTB7 292D 777 E. University Drive Tempe, AZ 85287-8404
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Mail code: 8404Campus: Tempe
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Ian Gilby has studied the behavioral ecology of wild chimpanzees since 1997. His main research interests are cooperative hunting, meat sharing and adult male dominance strategies. He is co-director of the Gombe Chimpanzee Database, which contains over five decades of detailed behavioral, ecological and demographic data from the long-term study of two chimpanzee communities in Gombe National Park, Tanzania. Before coming to Arizona State University in 2014, he was a postdoctoral research associate in human evolutionary biology at Harvard University, and then a senior research scientist in evolutionary anthropology at Duke University.
- Ph.D. Zoology, University of Minnesota 2004
- B.A. Biology, Carleton College 1996
Ian Gilby's research lies at the interface of behavioral ecology and evolutionary anthopology. The broad goal of his research program is to understand the roots of human cooperation using primate behavioral ecology. To date, his work has focused on hunting, meat sharing and social bonds among wild chimpanzees in Gombe National Park (Tanzania) and Kibale National Park (Uganda).
Lonsdorf, E. V., Gillespie, T.R., Wolf, T.M., Lipende, I., Raphael, J., Bakuza, J., Murray, C.M., Wilson, M.L., Kamenya, S., Mjungu, D., Collins, D.A., Gilby, I.C., Stanton, M.A., Terio, K.A., Barbian, H.J., Li, Y., Ramirez, M., Krupnick, A., Seidl, E., Goodall, J., Hahn, B.H., Pusey, A.E., Travis, D.A. 2017. Socioecological correlates of clinical signs in two communities of wild chimpanzees ( Pan troglodytes ) at Gombe National Park, Tanzania. In press, American Journal of Primatology. DOI: 10.1002/ajp.22562
Wood, B., Gilby, I.C. 2017. From Pan to man the hunter: hunting and meat sharing by chimpanzees, humans, and our common ancestor. In: Chimpanzees and Human Evolution (Ed. by Muller, M.N., Wrangham, R.W., & Pilbeam, D.). pp 339-382. Cambridge: Harvard University Press.
Gilby, I.C., Machanda, Z.P., O’Malley, R.C., Murray, C.M., Lonsdorf, E.V., Walker, K., Mjungu, D.C., Otali, E., Muller, M.N., Emery Thompson, M., Pusey, A.E., Wrangham, R.W. Predation by female chimpanzees: toward an understanding of sex differences in meat acquisition in the last common ancestor of Pan and Homo. Journal of Human Evolution 110: 82-94. DOI: 10.1016/j.jhevol.2017.06.015
Bray, J., Pusey, A.E., Gilby, I.C. 2016. Incomplete control and concessions explain mating skew in male chimpanzees. Proceedings of the Royal Society Series B 283: 20162071. DOI: 10.1098/rspb.2016.2071
Borries, C., Sandel, A.A., Koenig, A., Fernandez-Duque, E., Kamilar, J.M., Amoroso, C., Barton, R.A., Bray, J., Di Fiore, A., Gilby, I.C., Gordon, A.D., Mundry, R., Port, M., Powell, L., Pusey, A.E., Spriggs, A., Nunn, C.L. 2016.Transparency, usability, and reproducibility: guiding principles for improving comparative databases using primates as examples. Evolutionary Anthropology 25: 232-238.
Foerster, S., Franz, M., Murray, C.M., Gilby, I.C., Feldblum, J.T., Walker, K.K., Pusey, A.E. 2016. Chimpanzee females queue but males compete for social status. Scientific Reports 6: 35404.
Stevens, J.R., Marewski, J.N., Schooler, L.J., Gilby, I.C. 2016. Reflections of the social environment in chimpanzee memory: applying rational analysis beyond humans. Royal Society Open Science 3:160293.
O’Malley, R.C., Stanton, M.A., Gilby, I.C., Lonsdorf, E.V., Pusey, A.E., Markham, A.C., Murray, C.M. 2016. Reproductive state and rank influence patterns of meat consumption in wild female chimpanzees (Pan troglodytes schweinfurthii). Journal of Human Evolution 90: 16-28.
Gilby, I.C., Machanda, Z.P., Mjungu, D.C., Rosen, J., Muller, M.N., Pusey, A.E., Wrangham, R.W. 2015. Impact hunters catalyse cooperative hunting in two wild chimpanzee communities. Philosophical Transactions of the Royal Society Series B 370: 20150005.
Foerster, S., McLellan, K., Schroepfer-Walker, K., Murray, C. M., Krupenye, C., Gilby, I.C., Pusey, A. E. 2015. Social bonds in the dispersing sex: partner preferences among adult female chimpanzees. Animal Behaviour 105: 139-152.
Feldblum, J. T., Wroblewski, E. E., Rudicell, R. S., Hahn, B. H., Paiva, T., Cetinkaya-Rundel, M., Pusey, A. E., Gilby, I. C. 2014. Sexually coercive male chimpanzees sire more offspring. Current Biology 24: 2855-2860.
Wilson, M.L., Boesch, C., Fruth, B., Furuichi, T., Gilby, I.C. Hashimoto, C., Hobaiter, C., Hohmann, G., Itoh, N., Koops, K., Lloyd, J., Matsuzawa, T., Mitani, J.C., Mjungu, D.C., Morgan, D., Muller, M.N., Mundry, R., Nakamura, M., Pruetz, J., Pusey, A.E., Riedel, J., Sanz, C., Schel, A., Simmons, N., Waller, M., Watts, D.P., White, F.J., Wittig, R.M., Zuberbühler, K., Wrangham, R.W. 2014. Lethal aggression in Pan is better explained by adaptive strategies than human impacts. Nature 513: 414-417.
Tennie, C., O'Malley, R.C., Gilby, I. C. 2014. Why do chimpanzees hunt? Considering the benefits and costs of acquiring and consuming vertebrate versus invertebrate prey. Journal of Human Evolution 71: 38-45.
Machanda, Z.P., Gilby, I.C., Wrangham, R.W. 2014. Mutual grooming among adult male chimpanzees: The immediate investment hypothesis. Animal Behaviour 87: 165–174.
Miller, J.A., Pusey, A.E., Gilby, I.C., Schroepfer-Walker, K., Markham, A.C., Murray, C.M. 2014. Competing for space: female chimpanzees are more aggressive inside than outside their core areas. Animal Behaviour 87: 147-152.
Machanda, Z.P., Gilby, I.C., Wrangham R.W. 2013. Male-female association patterns among free-ranging chimpanzees (Pan troglodytes schweinfurthii). International Journal of Primatology 34: 917-938.
Gilby, I.C., Wilson, M.L., Pusey, A.E. 2013. Ecology rather than psychology explains co-occurrence of predation and border patrols in male chimpanzees. Animal Behaviour 86: 61-74.
Gilby, I.C., Brent, L.J.N., Wroblewski, E.E., Rudicell, R.S., Hahn, B.H., Goodall, J., Pusey, A.E. 2013. Fitness benefits of coalitionary aggression in male chimpanzees. Behavioral Ecology and Sociobiology 67: 373-381.
Crofoot, M.C., Gilby, I.C. 2012. Cheating monkeys undermine group strength in enemy territory. Proceedings of the National Academy of Sciences, USA 109: 501-505.
Gilby, I.C. 2012. Nonkin cooperation: reciprocity, markets and mutualism. In: Evolution of Primate Societies (Ed. by Mitani, J., Call, J., Kappeler, P., Palombit, R., & Silk, J.). pp 514-530. Chicago, IL: University of Chicago Press.
Pintea, L., Pusey, A.E., Wilson, M.L., Gilby, I.C., Collins, D.A., Kamenya, S., Goodall, J. 2012. Long-term ecological changes affecting the chimpanzees of Gombe National Park, Tanzania. In: Long Term Changes in Africa’s Rift Valley: impacts on biodiversity and ecosystems (Ed. by Plumptre, A.J.). pp 194-210. New York: Nova Science Publishers.
Lonsdorf, E.V., Murray, C.M., Lonsdorf, E.V., Travis, D.A., Gilby, I.C. Chosy, J., Pusey, A.E., Goodall, J. 2011. A retrospective analysis of factors correlated to chimpanzee (Pan troglodytes schweinfurthii) respiratory health at Gombe National Park. EcoHealth 8:26-35.
Gilby, I.C., Emery Thompson, M., Ruane, J., Wrangham, R.W. 2010. No evidence of short-term exchange of meat for sex among chimpanzees. Journal of Human Evolution 59: 44-53.
Gilby, I.C., Pokempner, A. A., Wrangham, R. W. 2010. A direct comparison of scan and focal sampling methods for measuring wild chimpanzee feeding behaviour. Folia Primatologica 81: 254-264.
Gilby, I.C., Connor, R.C. 2010. The role of intelligence in group hunting: Are chimpanzees different from other social predators? In: The Mind of the Chimpanzee: Ecological and Experimental Perspectives (Ed. by Lonsdorf, E. V., Ross, S. R. & Matsuzawa, T.). pp 220-233. Chicago, IL: University of Chicago Press.
Foster, M.W., Gilby, I.C., Murray, C.M., Johnson, A., Wroblewski, E.E., Pusey, A.E. 2009. Alpha male chimpanzee grooming patterns: implications for dominance ‘style’. American Journal of Primatology 71: 136-144.
Tennie, C., Gilby, I.C., Mundry, R. 2009. The meat-scrap hypothesis: small quantities of meat may promote cooperative hunting in wild chimpanzees. Behavioral Ecology and Sociobiology 63: 421-431.
Gilby, I.C., Wrangham, R.W. 2008. Association patterns among wild chimpanzees (Pan troglodytes schweinfurthii) reflect sex differences in cooperation. Behavioral Ecology and Sociobiology 62: 1831-1842.
Gilby, I.C., Eberly, L.E., Wrangham, R.W. 2008. Economic profitability of social predation among wild chimpanzees: individual variation promotes cooperation. Animal Behaviour 75: 351-360.
Crofoot, M.C., Gilby, I.C., Wikelski, M.C., Kays, R.W. 2008. Interaction location outweighs the competitive advantage of numerical superiority in Cebus capucinus intergroup contests. Proceedings of the National Academy of Sciences, USA 105: 577-581.
Murray C.M., Gilby I.C., Mane S.V., Pusey A.E. 2008 Adult male chimpanzees inherit maternal ranging patterns. Current Biology 18: 20-24.
Wrangham, R.W., Crofoot M.C., Lundy R., Gilby I.C. 2007. Use of overlap zones in group-living primates: a test of the risk hypothesis. Behaviour 144: 1599-1619.
Gilby, I.C., Wrangham, R.W. 2007. Risk-prone hunting by chimpanzees (Pan troglodytes schweinfurthii) increases during periods of high diet quality. Behavioral Ecology and Sociobiology 61: 1771-1779.
Emery Thompson, M., Kahlenberg, S.M., Gilby, I.C., Wrangham, R.W. 2007. Within-community variation in habitat quality accounts for variance in reproductive success among female chimpanzees at Kibale National Park, Uganda. Animal Behaviour 73: 501-512.
Gilby, I.C., Eberly, L.E., Pintea, L., Pusey, A.E. 2006. Ecological and social influences on the hunting behaviour of wild chimpanzees, Pan troglodytes schweinfurthii. Animal Behaviour 72: 169-180.
Gilby, I.C. 2006. Meat sharing among the Gombe chimpanzees: Harassment and reciprocal exchange. Animal Behaviour 71: 953-963.
Stevens, J.R. Gilby, I.C. 2004. A conceptual framework for non-kin food sharing: timing and currency of benefits. Animal Behaviour 67: 603-614.
Courses
2025 Spring
Course Number | Course Title |
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ASB 493 | Honors Thesis |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 592 | Research |
ASB 499 | Individualized Instruction |
ASB 590 | Reading and Conference |
ASB 790 | Reading and Conference |
ASB 492 | Honors Directed Study |
ASM 104 | Bones, Stones/Human Evolution |
ASB 795 | Continuing Registration |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASB 499 | Individualized Instruction |
ASB 493 | Honors Thesis |
ASB 492 | Honors Directed Study |
ASB 595 | Continuing Registration |
2024 Fall
Course Number | Course Title |
---|---|
ASB 790 | Reading and Conference |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 590 | Reading and Conference |
ASB 499 | Individualized Instruction |
ASB 492 | Honors Directed Study |
ASB 493 | Honors Thesis |
ASB 592 | Research |
ASB 795 | Continuing Registration |
ASB 499 | Individualized Instruction |
ASM 443 | Primatology |
ASB 492 | Honors Directed Study |
ASB 493 | Honors Thesis |
2024 Summer
Course Number | Course Title |
---|---|
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 590 | Reading and Conference |
ASB 499 | Individualized Instruction |
ASB 499 | Individualized Instruction |
2024 Spring
Course Number | Course Title |
---|---|
ASB 493 | Honors Thesis |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 592 | Research |
ASB 499 | Individualized Instruction |
ASB 590 | Reading and Conference |
ASB 790 | Reading and Conference |
ASB 492 | Honors Directed Study |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASB 499 | Individualized Instruction |
ASB 493 | Honors Thesis |
ASB 492 | Honors Directed Study |
ASM 443 | Primatology |
2023 Fall
Course Number | Course Title |
---|---|
ASB 790 | Reading and Conference |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 590 | Reading and Conference |
ASB 499 | Individualized Instruction |
ASB 492 | Honors Directed Study |
ASB 493 | Honors Thesis |
ASB 592 | Research |
ASB 499 | Individualized Instruction |
ASM 300 | Anthropological Sciences Sem |
ASB 492 | Honors Directed Study |
ASB 493 | Honors Thesis |
2023 Summer
Course Number | Course Title |
---|---|
ASB 590 | Reading and Conference |
ASB 799 | Dissertation |
ASB 792 | Research |
2023 Spring
Course Number | Course Title |
---|---|
ASB 493 | Honors Thesis |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 592 | Research |
ASB 499 | Individualized Instruction |
ASB 590 | Reading and Conference |
ASB 790 | Reading and Conference |
ASB 492 | Honors Directed Study |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASB 499 | Individualized Instruction |
ASB 493 | Honors Thesis |
ASB 492 | Honors Directed Study |
2022 Fall
Course Number | Course Title |
---|---|
ASB 790 | Reading and Conference |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 590 | Reading and Conference |
ASB 499 | Individualized Instruction |
ASB 492 | Honors Directed Study |
ASB 493 | Honors Thesis |
ASB 592 | Research |
ASB 499 | Individualized Instruction |
ASM 443 | Primatology |
ASB 492 | Honors Directed Study |
2022 Summer
Course Number | Course Title |
---|---|
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 590 | Reading and Conference |
2022 Spring
Course Number | Course Title |
---|---|
ASB 493 | Honors Thesis |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 592 | Research |
ASB 499 | Individualized Instruction |
ASB 590 | Reading and Conference |
ASB 790 | Reading and Conference |
ASB 492 | Honors Directed Study |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASB 499 | Individualized Instruction |
ASM 443 | Primatology |
2021 Fall
Course Number | Course Title |
---|---|
ASB 790 | Reading and Conference |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 590 | Reading and Conference |
ASB 499 | Individualized Instruction |
ASB 492 | Honors Directed Study |
ASB 493 | Honors Thesis |
ASB 592 | Research |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
2021 Summer
Course Number | Course Title |
---|---|
ASB 799 | Dissertation |
ASB 590 | Reading and Conference |
2021 Spring
Course Number | Course Title |
---|---|
ASB 493 | Honors Thesis |
ASB 499 | Individualized Instruction |
ASB 492 | Honors Directed Study |
ASB 584 | Internship |
2020 Fall
Course Number | Course Title |
---|---|
ASB 790 | Reading and Conference |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 590 | Reading and Conference |
ASB 499 | Individualized Instruction |
ASB 492 | Honors Directed Study |
ASB 493 | Honors Thesis |
ASB 592 | Research |
ASM 443 | Primatology |
2020 Summer
Course Number | Course Title |
---|---|
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 590 | Reading and Conference |
2020 Spring
Course Number | Course Title |
---|---|
ASB 493 | Honors Thesis |
ASB 799 | Dissertation |
ASB 792 | Research |
ASB 592 | Research |
ASB 499 | Individualized Instruction |
ASB 590 | Reading and Conference |
ASB 790 | Reading and Conference |
ASB 492 | Honors Directed Study |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASM 104 | Bones, Stones/Human Evolution |
ASB 499 | Individualized Instruction |
ASB 584 | Internship |
American Association of Physical Anthropologists International Primatological Society Sigma Xi, The Scientific Research Society
- SHESC Faculty Honors Advisor
- SHESC Undergraduate Researc Committee