Robert Page

Provost Emeritus and Provost Emeritus,

Office of University Provost

Other ASU affiliations

Biosocial Complexity Initiative

Faculty, Center for Social Dynamics and Complexity (CSDC)

School of Complex Adaptive Systems

Provost Emeritus; Foundation Chair of Life Sciences, School of Life Sciences

Emeritus Professor, Emeritus College

Affiliate Global Futures Scholar, Global Futures Scientists and Scholars

ISTB1-391F PO Box 874804 TEMPE, AZ 85287-6505
Mail code: 7805

Campus: Tempe
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Long Bio

Regents Professor Robert E. Page, Jr. is Provost Emeritus, a University Professor and Foundation Chair of Life Sciences. He is also a distinguished sustainability scholar in ASU's Julie Ann Wrigley Global Institute of Sustainability. He previously served as the vice provost and dean of the College of Liberal Arts and Sciences (2011-2013) and the founding director of the School of Life Sciences (2004-2011). He joined ASU in 2004 after spending 15 years on the faculty of the University of California at Davis where he served as chair of entomology from 1999-2004. His background is in behavior and population genetics and the focus of his current research is on the evolution of complex social behavior. Using the honey bee as a model, he has dissected their complex foraging division of labor at all levels of biological organization from gene networks to complex social interactions.

Professor Page has published more than 250 research papers and articles, five books, and is listed as a “highly-cited author” by the ISI Web of Knowledge, representing the top 1/2 of 1% of publishing scientists. He received the Alexander von Humboldt Senior Scientist Award (the Humboldt Prize) in 1995, the highest honor given by the German government to foreign scientists. In 2010 he was elected to the Leopoldina - the German National Academy of Science, the longest continuing academy in the world. He is also an elected foreign member of the Brazilian Academy of Science (1999), a fellow of the American Association for the Advancement of Science (1991), elected member of the American Academy of Arts and Sciences (2006), a fellow of the Wissenschaftskolleg zu Berlin (2009-2010), a fellow of the California Academy of Sciences (2016) and was named a Carl Friedrich von Siemens Foundation fellow (2017-2018). He received the James W. Creasman Award of Excellence from the Arizona State University Alumni Association in 2018.

Education

Ph.D. Entomology, University of California, Davis 1980
B.S. Entomology; Minor in Chemistry. San Jose State University 1976

Curriculum Vitae

Video

Research Interests

The focus of current research is on the evolution of complex social behavior. Using the honey bee as a model, Regents' Professor Page has dissected their complex foraging division of labor at all levels of biological organization from gene networks to complex social interactions.

Publications

Authored Books:

Page, R. E. 2013. The Spirit of the Hive: the mechanisms of social evolution. Harvard University Press, Cambridge.
Laidlaw, H. H. and R. E. Page. 1998. Queen Rearing and Bee Breeding. Wicwas Press.

Publications:

Y. Wang, J. B. Campbell, O. Kaftanoglu, R. E. Page, G. V. Amdam, J. F. Harrison. 2016. Larval starvation improves metabolic response to adult starvation in honey bees (Apis mellifera L.).Journal of Experimental Biology 219 (7), 960-968.
Y. Wang, O. Kaftanoglu, C. S. Brent, R. E. Page, G. V. 2016. Amdam. Starvation stress during larval development facilitates an adaptive response in adult worker honey bees (Apis mellifera L.).Journal of Experimental Biology 219 (7), 949-959.
D. A. Galbraith, Y. Wang, G. V. Amdam, R. E. Page, C. M. Grozinger. 2015. Reproductive physiology mediates honey bee (Apis mellifera) worker responses to social cues. Behavioral ecology and sociobiology 69 (9), 1511-1518.
K.. E. Ihle, O. Rueppell, Z. Y. Huang, Y. Wang, M. K. Fondrk, R. E. Page Jr. 2015. Genetic architecture of a hormonal response to gene knockdown in honey bees. Journal of Heredity 106 (2), 155-165.
Kirsten S. Traynor, Yves Le Conte, and Robert E. Page Jr. 2015. Age matters: pheromone profiles of larvae differentially influence foraging behaviour in the honeybee, Apis mellifera. Animal Behaviour. 99: 1-8.
K..S. Traynor, Y. Le Conte, R. E. Page. 2014. Queen and young larval pheromones impact nursing and reproductive physiology of honey bee (Apis mellifera) workers.Behavioral ecology and sociobiology 68 (12), 2059-2073.
R. E. Page, E. R Nash, E. H. Erickson. 2014. Use of honey yield data to assess damage to the beekeeping industry of the United States. Bulletin of the Entomological Society of America 33 (3), 190-194.
Wang, Y., O. Kaftanoglu, M. K. Fondrk, R. E. Page. 2014. Nurse bee behaviour manipulates worker honeybee (Apis mellifera L.) reproductive development. Animal Behavior 92: 253-261.
Siegel A. J., M. K. Fondrk, G. V. Amdam and R. E. Page. 2013. In hive patterns of temporal polyethism in strains of honey bees (Apis mellifera) with distinct genetic backgrounds. Behav. Ecol. Sociobiol. 67: 1623-1632.
Siegel, A. J., C. Freedman, and R. E. Page. 2012. Ovarian control of nectar collection in the honey bee (Apis mellifera). PLoS One 7(4): e33465.
Wang, Y., S. D. Kocher, T. A. Linksvayer, C. M. Grozinger, R. E. Page, and G. V. Amdam. 2012. Regulation of behaviorally associated gene networks in worker honey bee ovaries. Journal of Experimental Biology 215: 124-134.
Siegel, A. J., O. Kaftanoglu, M. K. Fondrk, N. R. Smith, and R. E. Page. 2012. Ovarian regulation of foraging division of labor in Africanized backcross and pollen-hoarding honeybees. Animal Behavior 83: 653-658.
Leimar, O., K. Hartfelder, M. D. Laubichler, and R. E. Page. 2012. Development and evolution of caste dimorphism in honey bees -- a modeling approach. Ecology and Evolution 2: 3098-3109.
Page, R. E., M. K. Fondrk, and O. Rueppell. 2012. Complex pleiotropy characterizes the pollen hoarding syndrome in worker honey bees (Apis mellifera L.). Behavioral Ecology and Sociobiology 66: 1459-1466.
Ament, S. A., C. A. Blatti, C. Alaux, M. M. Wheeler, A. L. Toth, Y. Le Conte, G. J. Hunt, E. Guzmán-Novoa, G. DeGrandi-Hoffman, J. L. Uribe-Rubio, G. V. Amdam, R. E. Page, S. L. Rodriguez-Zas, G. E. Robinson, and S. Sinha. 2012. New meta-analysis tools reveal common transcriptional regulatory basis for multiple determinants of behavior. Proceedings of the National Academy of Science USA 109: E1801–E1810.
Strassmann, J. E., R. E. Page, G. E. Robinson, and T. D. Seeley. Brief Communication Arising: Kin selection and eusociality. 2011. Nature 471, doi:10.1038/nature09833.
Linksvayer, T. A., O. Kaftanoglu, E. Akyol, S. Blatch, G. V. Amdam, and R. E. Page. 2011. Larval and nurse worker control of developmental plasticity and the evolution of honey bee queen-worker dimorphism. Journal of Evolutionary Biology 24: 1939-1948.
Rueppell, O., J. D. Metheny, T. Linksvayer, M. K. Fondrk, R. E. Page, and G. V. Amdam. 2011. Genetic architecture of ovary size and asymmetry in European honeybee workers. Heredity 106: 894-903.
Kaftanoglu, O., T. A. Linksvayer, and R. E. Page. 2011. Rearing honey bees, Apis mellifera, in vitro 1: Effects of sugar concentrations on survival and development. Journal of Insect Science 11: article 96.
Graham, A. M., M. D. Munday, O. Kaftanoglu, R. E. Page, G. V. Amdam, and O. Rueppell. 2011. Support for the reproductive ground plan hypothesis of social evolution and major QTL for ovary traits of Africanized worker honey bees (Apis mellifera L.). BMC Evolutionary Biology 11:95.
Kaftanoglu, O., T. A. Linksvayer, and R. E. Page. 2010. Rearing honey bees (Apis mellifera L.) in vitro: effects of feeding intervals on survival and development. Journal of Apicultural Research 49: 311-317.
Sirviö, A., P. Pamilo, R. A. Johnson, R. E. Page, and J. Gadau. 2010. Origin and evolution of the dependent lineages in the genetic caste determination system of Pogonomyrmex ants. Evolution 65: 869-884.
Wang, Y., O. Kaftanoglu, A. Siegel, R. E. Page, and G. V. Amdam. 2010. Surgically increased ovarian mass in the honey bee confirms link between reproductive physiology and worker behavior. Journal of Insect Physiology 56: 1816-1824.
Ihle, K. E., R. E. Page, K. Frederick, M. K. Fondrk, and G. V. Amdam. 2010. Genotypic effect on regulation of behaviour by vitellogenin supports reproductive origin of honeybee foraging bias. Animal Behavior 79:1001-1006.
Amdam, G. V., R. E. Page, M. K. Fondrk, and C. S. Brent. 2010. Hormone response to bidirectional selection on social behavior. Evolution and Development 12: 428-436. [cover article]
Wang, Y., G. V. Amdam, M. A. Wallrichs, M. K. Fondrk, O. Kaftanoglu, and R. E. Page. 2009. PDK1 and HR46 gene homologs tie social behavior to ovary signals. PLoS One 4: e4899.
Amdam, G. V., O. Rueppell, M. K. Fondrk, R. E. Page, C. M. Nelson. 2009. The nurse’s load: early-life exposure to brood-rearing affects behavior and lifespan in honey bees (Apis mellifera). Experimental Gerontology 44: 467-471.
Rueppell, O., O. Kaftanoglu, and R. E. Page. 2009. Honey bee (Apis mellifera) workers live longer in small than in large colonies. Experimental Gerontology 44: 447-452.
Linksvayer, T. A., O. Rueppell, A. Siegel, O. Kaftanoglu, R. E. Page, and G. V. Amdam. 2009. The genetic basis of transgressive ovary size in honeybee workers. Genetics 2009 183: 693–707. [cover article]
Tsuruda, J. M. and R. E. Page. 2009. The effects of young brood on the foraging behavior of two strains of honey bees (Apis mellifera). Behavioral Ecology and Sociobiology 64: 161-167.
Tsuruda, J. M. and R. E. Page. 2009. The effects of foraging role and genotype on light and sucrose responsiveness in honey bees (Apis mellifera L.). Behavioural Brain Research 205: 132-137.
Amdam, G. V and R. E. Page. 2009. Oldroyd and Beekman do not test ground plan hypothesis that explains origins of social behavior. PLoS Biology 6, e56r2248.
Linksvayer, T. A. and R.E. Page 2009. Honey bee social regulatory networks are shaped by colony-level selection. The American Naturalist 173 (3) E-article
Al-Khafaji, K., S. Tuljapurkar, J. R. Carey, R. E. Page. 2009. Life in the colonies: Hierarchical demography of a social organism. Ecology 90: 556-566
Tsuruda, J. M., G. V. Amdam, and R. E. Page. 2008. Sensory response system of social behavior tied to female reproductive traits. PLoS One 3: e3397.
Page, R. E. 2008. Confessions of an evolutionary biologist. Biological Theory 2: 207-208. [Book review]
Rueppell, O., C. Bachelier, M. K. Fondrk, and R. E. Page. 2007. Regulation of life history regulates lifespan of worker honey bees. Experimental Gerontology 42: 1020-1032.
Page, R. E. and G. V. Amdam. 2007. The making of a social insect: developmental architectures of social design. Bioessays 29: 334-343. [cover article]
Nelson, C.M., K.E. Ihle, M.K. Fondrk, R.E. Page, G.V. Amdam. 2007. The gene vitellogenin has multiple coordinating effects on social organization. PloS Biology 5: 673-677. [With synopsis]
Rueppell, O., C. Bachelier, M. K. Fondrk, and R. E. Page. 2007. Regulation of life history regulates lifespan of worker honey bees. Experimental Gerontology 42: 1020-1032.
G. J. Hunt, G. V. Amdam, D. Schlipalius, C. Emore, N. Sardesai, C. E. Williams, O. Rueppell, E. Guzmán-Novoa, M. Arechavaleta-Velasco, S. Chandra, M. K. Fondrk, M. Beye, R. E. Page. 2007. Behavioral genomics of honeybee foraging and nest defense. Naturwissenschaften 94: 247-267. [cover article]
M. Beye, I. Gattermeier, M. Hasselmann, T. Gempe, M. Schioett, J. Baines, D. Schlipalius, F. Mougel, C. Emore, O. Rueppell, A. Sirviö, E. Guzmán-Novoa, G. Hunt, M. Solignac, R. E. Page. 2006. Exceptionally high levels of recombination across the honey bee genome. Genome Research 16: 1339-1344. [cover article]
The Honeybee Genome Sequencing Consortium. 2006. Insights into social insects from the genome of the honeybee Apis mellifera. Nature 443: 931-949. [cover article]
A Sirviö, J Gadau, O Rueppell, D Lamatsch, J Boomsma, P Pamilo, Robert Page. High recombination frequency creates genotypic diversity in colonies of the leaf-cutting ant Acromyrmex echinatior. Journal of Evolutionary Biology (2006).
G. Amdam, K. Norberg, Robert Page, J. Erber, R. Scheiner. Downregulation of vitellogenin gene activitiy increases the gustatory responsiveness of honey bee workers (Apis mellifera L.). Behavioral Brain Research (2006).
Gro Amdam, A. Csondes, M. Fondrk, Robert Page. Complex social behaviour derived from maternal reproductive traits. Nature (2006).
M Beye, O Rueppell, A Sirviö, E GuzmÃ¡n-Novoa, G. Hunt, M. Solignac, Robert Page, I Gattermeier, M. Hasselmann, T. Gempe, M. Schioett, J. Baines, D. Schlipalius, F. Mougel, C. Emore. Exceptionally high levels of recombination across the honey bee genome. Genome Research (2006).
O. Rueppell, S. Chandra, T. Pankiw, M. Fondrk, M. Beye, G. Hunt, Robert Page. The genetic architecture of sucrose responsiveness in the honey bee (Apis mellifera L.). Genetics (2006).
Robert Page. Insights into social insects from the genome of the honeybee Apis mellifera. Nature (2006).
Robert Page, R. Scheiner, J. Erber, G. Amdam. The development and evolution of division of labor and foraging specialization in a social insect. Current Topics in Developmental Biology (2006).
E. Guzmá¡n-Novoa, G. Hunt, Robert Page, J. Uribe-Rubio, D. Prieto-Merlos, F. Becerra-Guzman. Paternal effects on the defensive behavior of honeybees. Journal of Heredity (2005).
G. Amdam, Robert Page. Intergenerational transfers may have decoupled physiological and chronological age in a eusocial insect. Aging Research Reviews (2005).
M. Humphries, M. Fondrk, Robert Page. Locomotion and the pollen hoarding behavioral syndrome of the honey bee (Apis mellifera L.). Journal of Comparative Physiology A (2005).
O. Rueppell, M. Fondrk, Robert Page. Biodemographic analysis of male honey bee mortality. Aging Cell (2005).
O. Rueppell, Robert Page, M. Fondrk. Male maturation rate respond to selection on pollen hoarding in honeybees. Animal Behaviour (2005).
P. Hedrick, J. Gadau, Robert Page. Genetic sex determination and extinction. Trends in Ecology and Evolution (2005).
D. Schulz, T. Pankiw, K. Fondrk, G. Robinson, Robert Page. Comparisons of juvenile hormone hemolymph and octopamine brain titers in honey bees (Apis mellifera) selected for high and low pollen hoarding. Annals of the Entomological Society of America (2004).
G. Amdam, K. Norberg, M. Fondrk, Robert Page. Reproductive ground plan may mediate colony-level effects on individual foraging behavior in honey bees. Proceedings of the National Academy of Science USA (2004).
O. Rueppell, T. Pankiw, Robert Page. Pleiotropy, epistasis, and new QTL: the genetic architecture of honey bee foraging behavior. Journal of Heredity (2004).
O. Rueppell, G. Amdam, Robert Page, J. Carey. From genes to societies. Science of Aging Knowledge Environment (2004).
R. Scheiner, Robert Page, J Erber. Sucrose responsiveness and behavioral plasticity in honey bees. Apidologie (2004).
T. Pankiw, M. Nelson, Robert Page, M. Fondrk. The communal crop: modulation of sucrose response thresholds of pre-foraging honey bees with incoming nectar quality. Behavioral Ecology and Sociobiology (2004).
M. Beye, M. Hasselmann, M. Fondrk, Robert Page, S. Omholt. The gene csd is the primary signal for sexual development in the honeybee and encodes an SR-type protein. Cell (2003).
M Humphries, U MÃ¼ller, M Fondrk, Robert Page. PKA and PKC content in the honey bee central brain differs in genotypic strains with distinct foraging behavior. Journal of Comparative Physiology A (2003).
T Pankiw, Robert Page. The effect of pheromones, hormones, and handling on sucrose response thresholds of honey bees (Apis mellifera L.). Journal of Comparative Physiology A (2003).
D Tarpy, Robert Page. Sex determination and the evolution of polyandry in honey bees (Apis mellifera). Behavioral Ecology and Sociobiology (2002).
E GuzmÃ¡n-Novoa, G Hunt, Robert Page, M Fondrk. Genetic correlations among honey bee (Apis mellifera L.) behavioral characteristics and body size. Annals of the Entomological Society of America (2002).
G DeGrandi-Hoffman, Robert Page, J Martin, M Fondrk. Can the frequency of reduced Varroa jacobsoni (oud.) fecundity in honey bee (Apis mellifera L.) pupae be increased by selection?. Apidologie (2002).
J Gadau, Robert Page, J Werren. The genetic basis of interspecific differences in wing size in Nasonia (Hymenoptera: Pteromalidae): major quantitative trait loci and epistasis. Genetics (2002).
J Tomkins, Y Yu, G Hunt, R Wing, M Luo, G Fang, D Main, J Goicoechea, M Atkins, D Frisch, Robert Page, E GuzmÃ¡n-Novoa. New genomic resources for the honey bee (Apis mellifera L.): development of a deep-coverage BAC library and a preliminary STC database. Genetics and Molecular Research (2002).
Robert Page, J Erber. Levels of behavioral organization and the evolution of division of labor. Naturwissenschaften (2002).
Robert Page, Juergen Gadau, M Beye. The emergence of hymenopteran genetics. Genetics (2002).
T Pankiw, D Tarpy, Robert Page. Genotype and rearing environment affect honeybee perception and foraging behaviour. Animal Behavior (2002).
Guzmán-Novoa, E,Page, Robert Eugene. Successful selective breeding of honey bees (Apis mellifera L.) in an africanized area of Mexico. Proceedings of the 2nd International Conference on Africanized Honey Bees and Bee Mites (2002).
D Tarpy, Robert Page. The curious promiscuity of queen honey bees (Apis mellifera): evolutionary and behavioral mechanisms. Annales Zoologici Fennici (2001).
J Erber, Robert Page. The evolutionary dynamics of social organization in insect societies: from behavior to genes and back. Santa Fe Institute Bulletin 16 (2001).
J Gadau, C Gerloffs, N KrÃ¼gers, H Chan, P Schmid-Hempel, A Wille, Robert Page. A linkage analysis of sex determination in Bombus terrestris (L.) (Hymenoptera: Apidae). Heredity (2001).
M Hasselmann, M Fondrk, Robert Page, M Beye. Fine scale mapping in the sex locus region of the honey bee (Apis mellifera). Insect Molecular Biology (2001).
R Scheiner, Robert Page, J Erber. Responsiveness to sucrose affects tactile and olfactory learning in preforaging honey bees of two genetic strains. Behavioural Brain Research (2001).
R Scheiner, Robert Page, J Erber. The effects of genotype, foraging role and sucrose responsiveness on the tactile learning performance of honey bees (Apis mellifera L.). Neurobiology of Learning and Memory (2001).
Robert Page, C Peng. Aging and development in social insects with emphasis on the honey bee, Apis mellifera L. Experimental Gerontology (2001).
Robert Page, C Peng. Reply to letter to the editor. Experimental Gerontology (2001).
T Pankiw, K Waddington, Robert Page. Modulation of sucrose response thresholds in honey bees (Apis mellifera L.): influence of genotype, feeding, and foraging experience. Journal of Comparative Physiology A (2001).
T Pankiw, Robert Page. Brood pheromone modulates honey bee sucrose response thresholds (Apis mellifera L.). Behavioral Ecology and Sociobiology (2001).
T Pankiw, Robert Page. Genotype and colony environment affect honey bee (Apis mellifera L.) development and foraging behavior. Behavioral Ecology and Sociobiology (2001).
W Getz, Robert Page, P Starks. A tribute to William Donald Hamilton (1936-2000). Annales Zoologici Fennici (2001).
D Nielsen, P Ebert, Robert Page, G Hunt, E GuzmÃ¡n-Novoa. Improved polymerase chain reaction-based mitochondrial genotype assay for identification of the Africanized honey bee (Hymenoptera: Apidae). Annals of the Entomological Society of America (2000).
D Tarpy, Robert Page. No behavioral control over mating frequency in queen honey bees (Apis mellifera L.): implications for the evolution of extreme polyandry. The American Naturalist (2000).
E GuzmÃ¡n-Novoa, Robert Page. Pollen collecting and foraging force by European and European x Africanized hybrid honey bees (Hymenoptera: Apidae) in mixed genotype colonies are similar. Annals of the Entomological Society of America (2000).
J Fewell, Robert Page. Colony-level selection effects on individual and colony foraging task performance in honey bees, Apis mellifera L. Behavioral Ecology and Sociobiology (2000).
Juergen Gadau, Robert Page, J Werren, P Schmid-Hempel. Genome organization and social evolution in Hymenoptera. Naturwissenschaften (2000).
Robert Page, M Fondrk, G Hunt, E GuzmÃ¡n-Novoa, M Humphries, K Nguyen, A Green. Genetic dissection of honeybee (Apis mellifera L.) foraging behavior. Journal of Heredity (2000).
T Pankiw, Robert Page. Response thresholds to sucrose predict foraging division of labor in honeybees. Behavioral Ecology and Sociobiology (2000).

Research Activity

Page Jr,Robert Eugene*. Decoding a Pheromone Signal: How Brood Pheromone Stimulates Honey Bee Pollen Foraging Colony Health (Fellowship Kirsten Traynor). USDA-NIFA(8/15/2012 - 8/14/2014).
Laubichler,Manfred Dietrich*, Page Jr,Robert Eugene. Global Classroom - Liberal Arts Education for the 21st Century with International Perspective. Leuphana Univ(6/1/2012 - 12/31/2015).
Page Jr,Robert Eugene*, Amdam,Gro Vang. Caste Demography and Aging in Honey Bees. UNIV OF CA AT DAVIS(9/30/2009 - 8/31/2012).
Page Jr,Robert Eugene*. Postdoctoral Research Fellowship - TL. NSF-BIO(3/1/2006 - 2/28/2008).
Mchenry,Albert L*, Alford,Terry Lynn, Alford,Terry Lynn, Allison,Maria Teresa, Allison,Maria Teresa, Castillo-Chavez,Carlos, Crouch,Peter Evans, De Los Santos Jr,Alfredo G, Diaz,Rodolfo Enrique, Francisco,Wilson Alex, Garcia,Antonio Agustin, Garcia-Pichel,Ferran, Glick,Milton D, Johnson,Paul Carr, Keller,Gary D, Menendez,Jose, Moore,Ana L, Page Jr,Robert Eugene, Peralta,Pedro D, Phillips,Elizabeth Deutsch, Rivera,Daniel E, Rodriguez,Armando A, Villalobos,Jesus R, Wheeler,Quentin Duane, Young,David A. More Graduate Education at Mountain States Alliance (MGEMSA) AGEP Phase II (2004-2009). NSF-EHR(3/1/2005 - 3/31/2015).
Page Jr,Robert Eugene*. Pheromonal and Genetic Regulation of Honey Bee Foraging Behavior. USDA-CSREES(2/15/2005 - 2/14/2008).
Page Jr,Robert Eugene*. The Biodemography of Life Span. UNIV OF CA AT DAVIS(7/1/2004 - 6/30/2008).

Courses

2025 Spring

Course Number	Course Title
BIO 590	Reading and Conference
BIO 194	Special Topics
BIO 194	Special Topics

2024 Fall

Course Number	Course Title
BIO 194	Special Topics
BIO 194	Special Topics

2024 Spring

Course Number	Course Title
BIO 590	Reading and Conference
BIO 194	Special Topics
BIO 194	Special Topics

2023 Spring

Course Number	Course Title
BIO 590	Reading and Conference

2022 Spring

Course Number	Course Title
BIO 590	Reading and Conference

2021 Spring

Course Number	Course Title
BIO 590	Reading and Conference

2020 Spring

Course Number	Course Title
BIO 590	Reading and Conference

Presentations

Page, Robert. Genetics and neuroethology of honey bee foraging behavior. GDR 'd Ethologie (Nov 2005).
Page, Robert. Methods of breeding. French National Association of Queen Producers (Nov 2005).
Page, Robert. Selection for pollen collecting. French National Queen Producers (Nov 2005).
Page, Robert. The evolutionary origins of division of labor in the honey bee. 13th Altenberg Workshop on Theoretical Biology (Sep 2005).
Page, Robert. The genetic architecture of honey bee behavior - linkage mapping on a functional genome. 51st Congress of Brazilian genetics (Sep 2005).
Page, Robert. Evolutionary origins of division of labor in the honey bee. Sigma Xi Annual Banquet (May 2005).
Page, Robert. Social foraging in honey bees: genetics, neurobiology, and origins. Hexapodium, Center for Insect Science, University of Arizona (Feb 2005).
Page, Robert. European Section of the International Union for the Study of Social Insects

Honors / Awards

James W. Creasman Award of Excellence, 2018
Carl Friedrich von Siemens Foundation Fellowship, 2017
Elected Fellow, California Academy of Sciences, 2016
Regents' Professor, Arizona State University, 2015
Siemens Lecture – Carl Friedrich von Siemens Foundation, Munich, Germany, 2012
Elected Fellow, Entomological Society of America, 2012
Fellow, Wissenschaftskolleg zu Berlin, 2009-2010
German National Academy of Sciences - Leopoldina, Elected Full Member 2009
Ernst Mayr Lecture -- Berlin-Brandenburg Academy of Science, 2009
American Academy of Arts and Sciences, Elected Member 2006
ISI Highly Cited (Plant and Animal Science) 2005 – present [Represents the top 1/2 percent of publishing researchers.]
Brazilian Academy of Science, Elected Foreign Member, 1999
Alexander von Humboldt Senior Scientist Award, 1995
American Association for the Advancement of Science Fellow, 1992